How was it with your stock wheels?
I've owned both a WR-250R and a WR-250X, and both those bikes are like lots of bikes as far as this goes:
The right-hand spacer butts up against the head of the axle, not the right fork leg.
If the fork leg had to be squeezed in to contact the wheel spacer, the fork would bind.
The head of the axle contacts the right wheel spacer, which contacts the right wheel bearing inner race, which contacts the spacer in the center of the wheel, which contacts the left wheel bearing inner race, which contacts the left wheel spacer, which contacts the left fork leg.
Threading the axle nut onto the left end of the axle and tightening it pulls all of those part together under tension without pulling the right fork leg in and causing binding.
This is why you tighten the right side fork pinch bolts around the axle last.